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While the results were encouraging, by the mid-1990s Holst and Carolyn Deacon, a physiologist at the University of Copenhagen, confirmed an earlier finding by Rolf Mentlein, Baptist Galllwitz and Wolfgang Schmidt based at the University of Kiel in Germany who had found in 1993 that GLP-1 did not last long in the body because it was broken down in minutes by dipeptidyl peptidase-IV (DPP-4), an enzyme found in numerous tissues and in the bloodstream (Mentlein, Galllwitz, Schmidt

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Additionally, clinical trials with long-acting dual or triple agonists targeting the glucagon receptor have demonstrated a marked suppressive effect on circulating amino-acids in both humans and animals [66, 67]